Download
Software
Tools
Education
Standards
About NDB
A Portal for Three-dimensional Structural Information about Nucleic Acids
As of 8-Jul-2020 number of released structures: 10846
Search DNA
Search RNA
Advanced Search
Search for released structures
NDB ID:
1K4X
PDB ID:
1K4X
Nucleic Acid Backbone Torsions
CSV Format
Model ID
Chain ID
Residue Number
Residue Name
O3-P-O5-C5
P-O5-C5-C4
O5-C5-C4-C3
C5-C4-C3-O3
C4-C3-O3-P
C3-O3-P-O5
O4-C1-N1-9-C2-4
1
A
1
DG
0.0
0.0
182.84
172.25
183.06
256.79
113.97
1
A
2
DG
295.99
160.41
76.32
135.8
229.1
221.91
239.67
1
A
3
DG
356.68
203.94
303.9
160.55
216.17
209.58
47.02
1
A
4
DG
315.51
141.3
58.02
174.0
169.88
275.43
265.07
1
A
5
DT
269.09
161.74
83.39
97.95
210.83
228.38
226.62
1
A
6
DT
114.76
253.96
185.04
120.64
112.65
194.62
198.42
1
A
7
DT
325.53
149.32
39.15
150.4
191.57
300.63
176.69
1
A
8
DT
244.79
226.75
51.82
138.95
217.38
178.61
198.27
1
A
9
DG
258.25
31.67
177.0
118.92
177.12
271.16
38.74
1
A
10
DG
286.49
164.69
68.69
128.6
38.57
76.11
255.43
1
A
11
DG
60.67
115.35
181.38
144.81
84.09
54.98
68.62
1
A
12
DG
189.4
164.58
69.88
119.48
0.0
0.0
225.79
1
B
1
DG
0.0
0.0
182.8
172.22
183.11
256.76
113.96
1
B
2
DG
296.05
160.44
76.28
135.78
229.1
222.0
239.68
1
B
3
DG
356.58
203.99
303.98
160.59
216.2
209.59
47.05
1
B
4
DG
315.52
141.36
57.95
174.07
169.91
275.33
265.11
1
B
5
DT
269.13
161.8
83.37
97.9
210.91
228.44
226.6
1
B
6
DT
114.84
253.77
185.16
120.83
112.43
194.8
198.41
1
B
7
DT
325.43
149.33
39.13
150.05
191.72
300.81
176.66
1
B
8
DT
244.81
226.67
51.98
138.68
217.52
178.6
198.68
1
B
9
DG
258.44
31.15
176.77
118.87
177.44
271.58
39.07
1
B
10
DG
286.73
164.78
68.07
129.14
38.54
76.21
256.21
1
B
11
DG
60.91
115.13
181.44
145.03
83.89
54.9
68.56
1
B
12
DG
189.12
164.66
68.91
120.26
0.0
0.0
225.61
2
A
1
DG
0.0
0.0
188.02
114.48
58.78
65.41
75.93
2
A
2
DG
198.79
161.79
69.25
112.16
294.59
127.49
260.8
2
A
3
DG
333.31
157.5
313.66
156.31
215.67
218.29
57.82
2
A
4
DG
303.68
151.2
61.53
172.21
181.15
266.31
259.28
2
A
5
DT
279.11
161.67
78.88
102.86
204.86
236.68
224.91
2
A
6
DT
123.52
224.68
188.91
138.83
295.58
115.73
204.01
2
A
7
DT
158.35
186.92
59.83
143.8
195.33
264.38
199.49
2
A
8
DT
275.36
220.5
58.76
162.78
237.03
69.53
207.5
2
A
9
DG
266.98
151.65
65.04
114.95
168.34
275.84
63.66
2
A
10
DG
292.27
155.26
74.36
122.15
183.36
217.51
233.26
2
A
11
DG
58.25
156.86
302.51
152.54
184.83
285.56
64.48
2
A
12
DG
278.38
175.19
66.93
126.36
0.0
0.0
244.11
2
B
1
DG
0.0
0.0
188.11
114.57
58.81
65.35
76.03
2
B
2
DG
198.81
161.83
69.19
112.17
294.68
127.47
260.74
2
B
3
DG
333.12
157.51
313.78
156.18
215.61
218.39
58.04
2
B
4
DG
303.75
151.12
61.55
172.14
181.15
266.32
259.17
2
B
5
DT
279.13
161.62
78.9
102.68
204.93
236.87
224.97
2
B
6
DT
123.59
224.64
188.96
138.8
295.5
115.66
203.92
2
B
7
DT
158.42
186.93
59.78
143.5
195.57
264.59
199.55
2
B
8
DT
275.41
220.54
58.65
162.69
237.21
69.46
208.02
2
B
9
DG
267.05
151.7
64.76
114.86
168.65
276.1
64.03
2
B
10
DG
292.31
155.22
74.22
122.7
183.34
217.27
234.2
2
B
11
DG
58.26
156.88
302.54
152.64
184.72
285.5
64.66
2
B
12
DG
278.41
175.27
65.47
127.12
0.0
0.0
244.14
3
A
1
DG
0.0
0.0
196.27
138.54
171.79
247.46
111.42
3
A
2
DG
304.7
162.26
70.67
142.44
194.93
213.9
244.78
3
A
3
DG
33.75
200.69
303.42
161.32
195.86
232.62
42.63
3
A
4
DG
304.71
155.26
64.39
175.04
173.06
274.18
267.65
3
A
5
DT
272.3
159.21
86.33
99.94
208.23
242.16
223.49
3
A
6
DT
127.18
233.68
180.48
122.25
86.57
241.86
198.74
3
A
7
DT
299.09
188.54
48.32
149.48
192.65
297.03
186.06
3
A
8
DT
254.94
226.56
51.92
142.18
195.56
78.91
202.0
3
A
9
DG
87.95
240.17
182.67
117.01
178.86
272.14
51.02
3
A
10
DG
286.14
154.03
76.5
124.03
185.33
211.49
236.28
3
A
11
DG
56.56
158.84
299.93
156.47
173.88
287.89
69.17
3
A
12
DG
256.94
187.13
58.75
140.5
0.0
0.0
237.83
3
B
1
DG
0.0
0.0
196.24
138.61
171.74
247.5
111.45
3
B
2
DG
304.62
162.22
70.79
142.41
194.89
213.98
244.86
3
B
3
DG
33.74
200.7
303.49
161.28
195.83
232.64
42.73
3
B
4
DG
304.66
155.23
64.47
174.99
173.04
274.18
267.7
3
B
5
DT
272.29
159.17
86.32
99.97
208.17
242.19
223.54
3
B
6
DT
127.16
233.67
180.5
122.21
86.71
241.75
198.77
3
B
7
DT
299.21
188.51
48.19
149.2
192.83
297.27
185.87
3
B
8
DT
254.98
226.44
51.89
141.97
195.66
79.09
202.35
3
B
9
DG
87.73
239.93
182.45
116.9
179.17
272.55
51.49
3
B
10
DG
286.18
153.91
76.3
124.39
185.29
211.38
236.93
3
B
11
DG
56.72
158.7
300.06
156.41
173.75
287.86
69.14
3
B
12
DG
256.78
187.06
57.85
140.92
0.0
0.0
237.78
4
A
1
DG
0.0
0.0
181.4
138.74
171.16
257.26
107.37
4
A
2
DG
301.64
161.71
78.17
138.18
217.71
245.1
240.98
4
A
3
DG
335.65
208.98
310.04
165.75
215.15
201.2
62.2
4
A
4
DG
316.3
141.91
57.3
173.08
176.99
269.59
265.87
4
A
5
DT
280.77
154.96
80.5
101.96
203.71
233.38
228.34
4
A
6
DT
128.37
240.89
180.52
121.4
70.13
258.21
195.82
4
A
7
DT
279.86
209.78
53.35
147.95
188.33
292.48
190.94
4
A
8
DT
211.21
273.34
50.27
146.2
281.39
71.13
204.6
4
A
9
DG
186.98
97.34
171.85
112.47
59.99
86.02
46.46
4
A
10
DG
190.53
135.96
81.89
125.72
302.78
164.31
246.28
4
A
11
DG
318.3
194.43
293.89
149.36
174.15
284.41
72.64
4
A
12
DG
261.6
182.49
67.92
123.95
0.0
0.0
216.6
4
B
1
DG
0.0
0.0
181.38
138.75
171.13
257.37
107.35
4
B
2
DG
301.66
161.71
78.21
138.28
217.63
245.09
240.97
4
B
3
DG
335.58
209.02
310.12
165.64
214.97
201.25
62.32
4
B
4
DG
316.49
141.82
57.25
173.11
177.03
269.59
266.01
4
B
5
DT
280.69
155.02
80.55
101.88
203.69
233.42
228.39
4
B
6
DT
128.38
240.88
180.54
121.42
70.14
258.11
195.85
4
B
7
DT
279.93
209.69
53.37
147.58
188.49
292.72
190.93
4
B
8
DT
211.13
273.43
50.1
146.03
281.81
70.81
204.97
4
B
9
DG
187.03
96.96
171.73
112.31
60.61
85.96
46.88
4
B
10
DG
190.52
135.92
81.51
126.2
302.82
164.32
247.05
4
B
11
DG
318.14
194.21
294.39
149.27
174.04
284.2
72.61
4
B
12
DG
261.58
182.41
66.73
124.75
0.0
0.0
216.52
5
A
1
DG
0.0
0.0
35.89
140.35
165.34
264.63
95.41
5
A
2
DG
293.29
168.45
74.46
139.94
192.27
214.08
245.01
5
A
3
DG
35.77
195.95
305.14
156.76
198.52
239.93
38.53
5
A
4
DG
305.19
156.81
62.81
158.42
60.03
73.77
267.19
5
A
5
DT
198.88
151.08
85.5
93.75
196.02
230.47
227.03
5
A
6
DT
122.9
245.46
183.49
121.17
32.85
189.39
202.48
5
A
7
DT
115.02
68.41
37.48
130.18
195.51
283.66
185.18
5
A
8
DT
267.98
228.06
54.47
142.23
208.02
80.38
206.88
5
A
9
DG
89.35
223.2
181.86
120.05
165.76
275.1
49.86
5
A
10
DG
289.53
163.4
71.24
132.42
39.47
85.05
244.46
5
A
11
DG
55.26
127.25
180.61
159.27
181.65
281.9
63.74
5
A
12
DG
274.47
177.85
66.07
123.06
0.0
0.0
229.02
5
B
1
DG
0.0
0.0
35.87
140.39
165.38
264.54
95.39
5
B
2
DG
293.35
168.46
74.41
139.88
192.27
214.21
245.02
5
B
3
DG
35.69
196.02
305.18
156.71
198.57
239.91
38.5
5
B
4
DG
305.23
156.82
62.78
158.42
60.06
73.67
267.08
5
B
5
DT
198.88
151.15
85.44
93.8
196.01
230.47
226.87
5
B
6
DT
122.76
245.53
183.56
121.17
32.85
189.37
202.55
5
B
7
DT
115.08
68.34
37.45
129.78
195.76
284.01
185.06
5
B
8
DT
267.88
228.01
54.53
141.97
208.2
80.54
207.25
5
B
9
DG
89.12
222.92
181.67
120.07
166.0
275.49
50.6
5
B
10
DG
289.61
163.31
70.94
132.76
39.54
85.1
245.3
5
B
11
DG
55.49
127.21
180.45
159.14
181.53
281.96
63.66
5
B
12
DG
274.33
177.92
64.94
123.83
0.0
0.0
228.88
6
A
1
DG
0.0
0.0
309.68
136.39
164.86
258.94
99.67
6
A
2
DG
299.78
168.01
76.14
144.65
219.11
248.29
242.39
6
A
3
DG
331.11
210.38
309.87
165.8
198.27
212.32
66.69
6
A
4
DG
317.6
143.41
64.05
174.8
175.16
276.23
260.21
6
A
5
DT
276.23
155.08
82.46
98.51
210.1
233.02
228.63
6
A
6
DT
123.48
237.1
183.01
125.44
107.1
222.63
199.26
6
A
7
DT
304.01
173.02
38.21
144.21
196.02
293.39
183.37
6
A
8
DT
240.6
241.76
51.63
146.13
232.45
59.15
201.92
6
A
9
DG
96.96
211.04
186.19
109.6
54.69
90.7
53.56
6
A
10
DG
197.24
133.72
81.04
123.09
298.22
166.77
239.29
6
A
11
DG
323.73
195.05
300.86
137.53
86.81
57.81
66.12
6
A
12
DG
196.12
159.93
69.65
101.37
0.0
0.0
229.65
6
B
1
DG
0.0
0.0
309.34
136.57
164.87
258.94
99.85
6
B
2
DG
299.65
168.04
76.2
144.67
219.16
248.33
242.21
6
B
3
DG
330.99
210.48
309.76
165.86
198.31
212.35
66.55
6
B
4
DG
317.64
143.38
64.03
174.79
175.1
276.27
260.2
6
B
5
DT
276.27
155.14
82.41
98.46
210.04
233.08
228.69
6
B
6
DT
123.49
237.1
182.99
125.4
107.09
222.75
199.17
6
B
7
DT
303.88
173.02
38.11
143.72
196.32
293.69
183.54
6
B
8
DT
240.37
241.89
51.57
145.78
232.77
59.24
202.25
6
B
9
DG
96.67
210.89
186.17
109.27
55.28
90.75
54.15
6
B
10
DG
197.3
133.7
80.71
123.59
297.83
167.1
240.02
6
B
11
DG
323.5
195.0
301.23
137.42
86.55
57.93
66.1
6
B
12
DG
195.88
159.87
69.06
101.72
0.0
0.0
229.77
7
A
1
DG
0.0
0.0
316.56
116.36
66.84
63.6
73.73
7
A
2
DG
198.68
157.44
70.67
120.81
289.93
160.74
251.5
7
A
3
DG
332.69
197.58
301.61
154.47
205.81
233.79
43.41
7
A
4
DG
304.55
155.12
59.72
168.14
182.26
269.1
265.1
7
A
5
DT
283.04
157.91
79.52
89.76
197.18
245.01
214.94
7
A
6
DT
130.6
227.6
183.14
141.51
293.04
144.63
203.56
7
A
7
DT
182.19
148.19
49.71
128.06
85.08
72.6
193.37
7
A
8
DT
161.68
212.12
50.73
136.55
284.48
126.09
205.28
7
A
9
DG
204.62
47.24
173.03
107.31
62.03
79.98
44.58
7
A
10
DG
186.85
141.18
87.44
119.38
179.71
214.84
241.72
7
A
11
DG
57.34
161.0
303.99
140.71
79.34
61.26
64.75
7
A
12
DG
188.38
164.07
70.29
122.09
0.0
0.0
234.42
7
B
1
DG
0.0
0.0
316.64
116.33
66.84
63.57
73.81
7
B
2
DG
198.78
157.52
70.69
120.69
290.11
160.46
251.58
7
B
3
DG
332.81
197.48
301.67
154.23
205.63
233.99
43.86
7
B
4
DG
304.48
155.09
59.82
168.22
182.35
268.86
265.29
7
B
5
DT
283.26
157.89
79.44
89.47
197.15
245.32
214.77
7
B
6
DT
130.73
227.6
183.07
141.57
293.04
144.67
203.66
7
B
7
DT
182.15
148.07
49.48
127.73
85.39
72.91
193.49
7
B
8
DT
161.63
212.06
50.75
136.33
284.92
125.91
205.69
7
B
9
DG
204.79
46.7
172.97
107.17
62.76
79.71
44.96
7
B
10
DG
187.07
141.16
87.09
119.84
179.67
214.52
242.48
7
B
11
DG
57.49
160.83
304.17
140.52
79.24
61.31
64.71
7
B
12
DG
188.24
163.89
69.29
122.72
0.0
0.0
234.45
8
A
1
DG
0.0
0.0
39.38
133.57
160.69
256.01
101.8
8
A
2
DG
302.38
163.88
78.64
146.64
192.09
212.88
242.59
8
A
3
DG
39.89
179.42
300.91
159.31
350.14
144.83
49.2
8
A
4
DG
218.81
145.65
69.23
171.45
175.04
260.16
270.74
8
A
5
DT
286.53
158.66
80.96
102.39
198.24
245.88
218.88
8
A
6
DT
132.4
231.0
184.07
117.44
105.97
218.49
201.91
8
A
7
DT
301.62
174.19
41.13
148.19
192.03
294.13
182.07
8
A
8
DT
255.95
229.02
52.91
143.36
262.68
146.53
205.87
8
A
9
DG
203.71
52.62
165.71
121.07
171.18
275.91
39.51
8
A
10
DG
284.89
165.59
68.42
127.92
39.22
71.37
250.31
8
A
11
DG
68.97
118.72
183.5
160.53
182.08
287.98
64.09
8
A
12
DG
263.94
178.92
66.72
111.13
0.0
0.0
216.18
8
B
1
DG
0.0
0.0
39.35
133.65
160.8
255.99
101.84
8
B
2
DG
302.32
164.0
78.66
146.68
192.02
212.9
242.48
8
B
3
DG
39.94
179.52
300.82
159.37
350.09
144.87
49.13
8
B
4
DG
218.88
145.71
69.07
171.52
175.09
260.12
270.72
8
B
5
DT
286.65
158.63
80.89
102.35
198.2
245.85
218.78
8
B
6
DT
132.53
230.97
184.05
117.38
105.94
218.55
202.05
8
B
7
DT
301.69
174.19
40.96
147.86
192.33
294.33
181.87
8
B
8
DT
255.87
228.98
52.94
143.14
262.99
146.35
206.38
8
B
9
DG
203.89
52.07
165.49
120.87
171.5
276.58
39.9
8
B
10
DG
284.92
165.61
67.99
128.28
39.23
71.46
250.91
8
B
11
DG
69.03
118.74
183.51
160.38
182.0
288.02
63.99
8
B
12
DG
263.8
178.96
65.56
112.01
0.0
0.0
216.12
