A Portal for Three-dimensional Structural Information about Nucleic Acids
As of 13-Oct-2021 number of released structures: 11721
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NDB ID: 4U4R  PDB ID: 4U4R 


Base Pair Morphology Step Parameters

CSV Format

Model NumberStep NumberStep NameShiftSlideRiseTiltRollTwistX-DisplacementY-DisplacementHelical RiseInclinationTipHelical Twist
14022_A39A40:G467C469_221.506-2.3513.425-4.833-11.413-9.02516.1362.550.74849.793-21.088-15.321
14122_A40A41:U466G467_220.153-0.3453.632-5.96611.23464.733-0.821-0.4083.51310.3875.51765.837
14222_A41A438:G465U466_220.3460.053.565-1.3552.65573.181-0.051-0.3383.5592.2261.13673.233
15322_A46G48:C431G432_22-2.639-2.5492.26168.282-130.4497.932-0.5351.7412.398-68.657-35.94158.656
15722_A51G53:C427U440_225.231-1.0025.953-6.0943.174171.818-0.508-2.6335.9311.5913.055171.832
16322_A67G69:U82G83_220.762-0.4043.648-2.5585.25161.729-0.661-0.8713.5755.1062.48761.978
16522_A93A399:U395G396_22-1.868-0.1722.96-5.1294.231-5.091-6.788-22.4530.744-33.469-40.575-8.372
18322_A103C309:G357U358_221.28-2.2213.227-6.6613.90190.248-1.639-1.0293.0712.7494.69390.507
18222_A105A103:U358C308_223.713-4.837-2.19243.45692.43622.61-0.644-3.498-3.25573.306-34.463103.926
19022_A108G109:C305U306_22-0.205-0.9763.148-3.62110.76930.379-3.493-0.222.66319.7056.62632.387
19422_A112U113:A301U302_22-1.07-1.3553.096-2.69313.30724.07-5.641.6772.16629.1315.89627.586
19722_A119U120:U297C298_222.923-1.5923.253-13.5163.39446.015-2.188-4.5132.2434.22916.84147.969
110222_A124U125:U292U293_220.506-1.093.6542.0648.54243.896-2.283-0.4593.41411.291-2.72944.724
110922_A145U146:A168A169_22-0.07-0.7763.198-10.235-6.59832.722-0.287-1.4483.17-11.23717.43334.857
111122_A147A148:C166U167_220.993-1.9733.9-0.36711.44345.573-3.542-1.2823.33214.5040.46646.915
111222_A148C149:G165C166_223.804-2.8543.699-4.74919.47942.34-5.068-5.161.86725.3396.17846.65
111822_A182U183:A202U203_22-0.238-1.1852.8-0.42213.19724.843-4.8340.4141.93728.2690.90528.085
112422_A188C189:G196A197_220.9-1.2113.611.454-3.25255.936-1.081-0.8663.69-3.463-1.54856.04
112922_A206U207:C258U259_220.658-1.5153.6340.06117.14636.874-4.137-0.9432.725.487-0.09140.54
113322_A210U211:A254U255_220.908-1.7443.5093.5434.63730.543-4.187-0.9823.2968.702-6.64931.083
113622_A213G214:A251U252_22-0.856-0.7873.285-3.86211.26460.441-1.2960.6573.15211.0733.79761.493
114022_A236C237:G228U229_220.623-0.863.292-3.17612.24647.816-1.916-0.9752.95614.8083.84149.367
115222_A265U267:A288U289_22-1.74-0.7962.7740.7773.64443.274-1.3712.4142.6714.93-1.05243.427
110522_A266G142:A173U174_221.897-2.6073.5981.651-3.6180.565-1.909-1.4183.724-2.79-1.27680.646
18822_A312A315:A352A353_225.148-0.5473.1396.502-21.7724.8699.692-9.4852.615-74.798-22.33723.231
116122_A316C317:G347U348_220.142-2.0523.3223.2733.74330.26-4.6050.3643.057.11-6.21630.657
116422_A323U324:A344U345_22-1.166-1.0853.127-8.97817.87240.118-2.9570.7672.63324.31312.21344.643
116922_A328G329:C339U340_220.35-1.6473.3852.5237.8930.377-4.48-0.1852.89514.716-4.70531.461
117222_A366A367:U374U375_22-0.111-1.5622.986-1.894.18532.394-3.413-0.0912.7697.4543.36732.709
117322_A367U368:G373U374_220.307-1.3972.9171.8787.25539.504-2.741-0.2652.64110.617-2.74840.18
16622_A399A401:C394U395_22-0.032-1.6263.065-6.663-1.9337.289-2.269-0.7623.103-2.98610.31137.906
16722_A401G404:C393C394_223.247-1.2833.6022.464-0.63178.777-0.991-2.4823.694-0.497-1.94178.811
17122_A407C408:G389G390_22-1.3011.4412.9132.7159.105-15.771-8.559-2.8621.982-29.9038.918-18.396
17422_A410C411:G422G423_220.94-1.0623.004-10.1163.78525.479-2.999-4.0432.2928.13121.73427.639
14322_A438U439:A464G465_22-3.614-3.5122.63918.8015.25210.618-9.1349.428-2.61316.607-59.4522.197
14522_A441C442:G462U463_22-0.82-2.5953.316-0.193.43531.738-5.3261.4573.0316.2580.34631.919
16122_A452G63:C87U88_22-2.896-1.7293.1183.6222.68851.995-2.1283.5122.8383.06-4.12352.177
1122_A468G595:C31U32_221.417-2.8373.694-4.7737.80883.501-2.323-1.1833.3985.8473.57483.909
117722_A477A478:G510A511_22-0.624-2.4993.459-1.5144.36928.683-5.9490.913.088.7463.03129.045
117822_A478C479:G509G510_220.403-1.2822.751-2.25-5.8827.405-1.429-1.2932.915-12.2074.6728.105
118122_A481A483:U504A505_22-1.184-4.2155.09316.415.48844.033-6.5813.032.9519.234-20.36749.217
118222_A483A485:U502U504_220.298-3.5596.167-9.759-1.1660.81-3.376-1.1586.12-1.1389.57461.524
118322_A485G486:U501U502_222.064-2.3334.86116.2411.69122.362-8.2371.1693.85724.711-34.32729.923
118622_A515G516:C536G537_221.124-0.5123.654-5.1655.39556.365-0.867-1.4953.4895.6845.44256.818
118822_A518A520:U532U533_22-1.944-1.4873.7928.24520.95446.078-3.2292.8492.57225.088-9.87251.015
118922_A520U522:C531U532_22-0.285-1.5675.5515.43411.43757.51-2.4851.5334.97211.539-15.57160.373
119522_A550G557:C587U588_22-1.382-0.8273.0760.5360.45347.121-1.071.773.0530.567-0.6747.126
1622_A599U600:A26U27_220.776-1.0522.9813.5755.11631.797-2.697-0.8252.859.226-6.44732.388
1822_A601U602:G23U24_22-0.129-1.0793.315-0.48510.90439.266-2.7270.1342.92615.8570.70540.697
11122_A604C4:G20U21_22-2.683-1.1435.49134.6921.16857.6-1.1134.5353.5291.11-32.95466.474
120522_A615G616:C1105U1106_22-0.142-1.9743.5392.4028.95234.126-4.5850.5922.92814.916-4.00135.326
120722_A622G1032:U1103U1104_221.134-3.1513.1137.4577.1827.667-20.0711.1950.75837.163-38.58912.876
127122_A746C747:G802U805_223.617-0.8534.571-31.784.35861.059-0.953-4.6282.5694.00929.23368.245
127822_A760G761:A789U790_22-0.627-0.4623.621-0.7570.45164.089-0.4570.5553.6250.4250.71364.094
128022_A774G775:U785C786_221.211-4.98-0.238132.959-116.239-4.4571.087-1.0-4.19559.19367.707-176.609
128322_A814G816:A855U857_224.7140.7350.206-119.001111.47146.2250.39-2.335-0.64755.98159.764175.094
122522_A865G866:C962U965_222.89-1.6663.2032.0855.46648.159-2.438-3.3633.1236.669-2.54548.492
122922_A869C870:G957U958_220.147-1.0783.3420.9968.27145.222-2.092-0.1033.11110.649-1.28345.944
124022_A881U882:U946U947_220.631-1.0142.687-0.146-1.75440.732-1.296-0.9192.724-2.5190.2140.768
124322_A884G885:C927U928_22-0.442-1.0133.3875.7774.18135.481-2.2261.5323.1436.776-9.36336.168
124622_A887U888:A924G925_22-0.227-0.8153.272-4.3995.90933.477-2.299-0.2973.09210.1067.52534.256
129222_A900G902:A907U909_22-1.062-0.7626.64212.7818.44297.171-1.0981.1126.33812.176-8.43899.088
129322_A933U935:G876A944_220.004-3.046-0.222150.69680.495-143.4891.5230.0030.236-40.35575.55-177.136
121222_A966A967:U632U633_22-1.419-1.9933.147-2.0385.41434.692-4.0512.0682.8899.0013.38835.156
121322_A967U968:G631U632_220.302-1.1933.2560.4616.137.336-2.6-0.4083.0349.45-0.71537.816
121622_A970A971:G628U629_225.781-1.5443.221-0.2041.39636.55-2.648-9.2393.132.2250.32636.577
121722_A971G972:C627G628_220.741-0.6123.747-0.62512.74362.146-1.219-0.7373.56812.2070.59963.314
121922_A973A974:C625U626_22-0.043-1.3792.754-10.8593.94626.469-3.488-1.8932.3628.14222.40628.84
122022_A974C975:G624C625_220.104-1.4223.3062.50.60543.605-1.970.1023.2880.813-3.36243.677
129722_A979G980:C1021C1022_22-0.22-0.8973.7-3.36611.80649.787-1.94-0.0023.42513.7833.9351.185
130122_A983G984:U1017U1018_220.393-1.9922.9970.1412.57418.354-8.644-0.9821.36634.636-0.38522.218
130622_A992A993:G1011U1012_22-3.5430.5934.681-117.367-125.114-65.7110.806-2.810.87363.965-60.004-172.901
130722_A993G994:C1010G1011_220.135-0.7933.257-6.1568.42157.968-1.234-0.4483.0978.6136.29658.819
130922_A995U996:G1008U1009_220.412-1.9263.469-0.2723.58236.538-3.564-0.6933.2695.6970.43336.708
131122_A1036C1037:G1094U1095_220.7-1.6573.2285.7041.75635.405-2.939-0.3273.2152.862-9.29735.889
131322_A1061U1063:U1056U1058_226.134-3.0820.76562.087138.045-68.8231.2253.290.682-74.59533.55-156.455
131622_A1065C1066:G1053U1054_220.845-1.9913.3542.4332.10639.069-3.224-0.9633.2913.142-3.6339.196
132022_A1069C1070:G1048U1049_220.543-1.9893.2162.4017.32227.234-5.621-0.6032.63915.168-4.97328.284
132722_A1076C1077:G1041G1042_221.104-1.2432.7425.4821.60124.21-3.302-1.1412.8323.751-12.84424.865
133722_A1113U1115:A1131A1132_222.482-2.2541.419122.194131.357-121.2490.8741.4761.217-65.70761.123-179.708
133922_A1116U1117:U313G1130_22-3.45-1.9562.575-1.3449.84134.12-4.2555.5152.07816.3472.23235.495
11622_A1138U8:U15G16_22-1.0320.3032.917-3.7031.45711.702-0.280.5013.1066.89217.51412.358
12022_A1142A1143:A11U12_220.175-1.7363.6483.778.34634.844-4.0670.2863.16313.648-6.16535.991
12122_A1143U1144:G10A11_22-0.987-1.7963.386-2.0334.81932.188-4.0411.4043.1468.6213.63732.599
12522_A1147C1148:G1629U1630_220.263-1.2323.0655.4959.25129.765-3.7880.4242.58217.312-10.28431.609
12822_A1151A1152:U1626U1627_22-1.138-1.6833.581-0.239-1.18135.287-2.5821.8373.641-1.9470.39435.307
12922_A1152G1153:C1625U1626_220.08-1.8582.876-1.5913.8226.721-4.788-0.5112.5818.2023.41627.033
13422_A1157A1160:U1617C1618_223.604-1.172.751-10.811-4.64767.558-0.905-3.5172.287-4.1469.64768.456
13522_A1160C1161:G1616U1617_220.078-0.9893.4385.21-2.72932.232-1.2520.8193.478-4.864-9.28832.75
136622_A1203U1560:U1551G1553_221.0930.56-5.348-17.2225.752-58.077-0.1062.427-4.91-5.772-17.282-60.612
138022_A1211G1212:C1451U1452_221.084-1.7393.271-0.50110.2721.709-7.106-2.7512.20525.5061.24323.994
138522_A1219C1220:G1263G1264_220.649-0.5243.634-1.174-2.26166.915-0.376-0.643.637-2.0491.06566.958
138722_A1221C1222:G1261U1262_220.263-0.9373.0040.980.81631.424-1.868-0.3182.9861.505-1.80831.45
138922_A1223A1224:U1259U1260_220.192-1.8233.4167.00619.0936.435-4.4470.4192.23328.015-10.28141.557
139322_A1234G1245:U1250U1251_222.5421.815-0.63137.909-98.024-9.184-1.3670.635-1.06751.8973.003-169.232
139522_A1236G1237:C1248U1249_220.744-1.5023.5263.994.53630.771-3.686-0.5813.3438.446-7.42931.345
140322_A1275U1276:A1436C1431_220.53-4.153.4274.62417.184-23.112.782.535.068-36.7059.877-29.093
141922_A1296G1297:A1300U1301_22-4.152-1.823.3052.8913.35755.877-2.5664.4692.64314.022-3.03457.393
143122_A1344A1345:U1380U1381_22-0.71-0.6823.0151.709-2.02630.672-0.9191.6453.01-3.822-3.22430.783
143222_A1345A1348:U1377U1380_221.8521.7534.12321.7846.803101.2750.988-0.7624.4374.362-13.965103.121
143322_A1348G1349:C1376U1377_220.37-1.6153.279-6.07921.99424.76-5.753-1.4571.32741.70411.52633.55
144122_A1357G1358:C1365U1366_220.127-1.533.0597.22414.10929.713-4.5940.7762.11425.368-12.9933.592
144322_A1388C1389:G1412G1409_22-2.5635.552-2.769-11.91-6.57935.7867.7575.227-2.752-10.26118.57638.206
144522_A1391U1392:A1406U1407_22-0.561-1.5753.3710.9566.6735.058-3.5221.0523.01510.948-1.56935.68
136322_A1469C1470:G1574G1170_22-0.1940.1583.082-0.0265.47924.724-1.1470.4353.04512.5990.05925.315
145022_A1471C1472:G1536U1538_221.735-2.3993.636-6.38215.322-74.1851.4031.1834.112-12.551-5.228-75.758
145322_A1475C1476:G1531U1532_220.743-1.3333.1626.3643.68139.176-2.369-0.3793.1085.432-9.3939.833
145722_A1479G1480:C1527U1528_22-0.687-2.2933.8246.8948.39431.746-5.4452.3972.93114.812-12.16633.508
145922_A1487G1488:C1518U1519_220.465-1.4532.663-7.994-1.61325.424-2.808-2.6922.489-3.55117.59526.68
137622_A1569A1570:G1541G1542_22-0.375-0.0812.647-5.1494.83225.413-1.236-0.3052.6110.72311.42526.36
137722_A1570C1571:G1540G1541_220.764-0.5783.188-2.8539.37767.975-0.848-0.7793.0658.3422.53968.594
134522_A1576A1577:U1168G1169_221.664-1.1823.7250.01611.72857.495-1.885-1.73.44612.055-0.01658.579
134622_A1577U1578:G1167U1168_220.263-1.3653.3174.352-2.98843.971-1.5240.0693.408-3.974-5.78744.271
146922_A1586A1587:U1609G1610_22-0.44-0.7913.025-6.3123.88358.51-0.9880.1473.0013.9556.42958.937
147022_A1587G1588:U1608U1609_22-0.199-1.7693.677-1.12411.90136.402-4.2620.1562.97918.4491.74238.251
147522_A1592A1593:U1603U1604_220.63-1.0173.394-5.2916.72341.8-2.078-1.4023.1039.3057.32342.628
147622_A1593G1594:C1602U1603_22-0.808-1.6322.950.5643.87531.109-3.6541.5882.7187.19-1.04731.349
147922_A1719G1720:A1681U1682_220.589-1.4632.8186.01516.735105.216-1.104-0.32.66110.483-3.768106.265
148122_A1721A1722:G1679G1680_22-0.509-0.5792.934-4.4817.1331.349-2.1520.2112.78312.9078.11132.433
148222_A1722U1723:A1678G1679_221.679-1.1523.401-0.637.71563.163-1.437-1.6223.2417.3370.59963.587
148822_A1728C1729:G1672G1673_220.829-0.983.175-5.36.19330.098-2.911-2.4742.74111.6589.97731.157
149022_A1730A1731:G1670A1671_220.608-2.0743.2136.911-5.163-22.4876.4453.4292.39412.65616.941-24.065
149122_A1731A1732:U1669G1670_220.436-0.7333.456-2.9019.34959.483-1.208-0.5833.2959.3532.90360.21
149222_A1732C1733:G1668U1669_22-0.625-1.2973.190.785.66934.987-2.9161.1352.9369.353-1.28735.437
150022_A1740U1741:A1660U1661_220.345-1.1022.9595.5355.64732.8-2.7140.2022.7569.824-9.62833.714
150422_A1744G1745:A1655U1656_22-1.772-0.6523.5253.86317.51360.564-1.4431.8783.14316.968-3.74362.921
150622_A1746G1747:C1653G1654_221.799-0.3443.993-5.58813.87360.444-1.095-2.0513.6813.5575.46162.096
150922_A1749A1750:U1650A1651_22-0.828-1.9013.164.6161.94820.869-5.8273.9292.7275.28-12.51321.456
151022_A1750C1751:G1649U1650_220.97-1.5673.186-0.4778.8829.001-4.615-1.9432.58817.2270.92630.306
151322_A1753A1754:C1646U1647_22-0.684-3.457-0.967-151.44684.463103.224-1.5260.706-1.43242.57876.344175.907
151422_A1754A1756:G1645C1646_22-2.84-2.5172.05988.285-104.707162.158-1.1461.5192.051-52.534-44.295173.478
151522_A1756G1757:C1644G1645_220.135-2.2353.484-5.73519.95938.261-4.795-0.6922.08428.0748.06643.347
152022_A1762A1763:C1639C1640_22-1.407-1.1993.596-4.963-5.59643.135-0.9971.3453.846-7.5436.6943.748
152122_A1763C1764:G1638C1639_22-0.761-1.3433.1516.1582.06340.784-2.1131.72.9422.937-8.76941.276
153022_A1776G1777:C1784U1785_22-0.349-1.0583.2-0.2785.91835.993-2.4730.5212.9979.4980.44736.462
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1225788_A43A44:U100U101_88-0.278-1.8513.5938.6431.89836.416-3.1521.6553.3452.984-13.58537.44
1225888_A44C45:G58U100_880.478-1.0612.997-4.1461.96643.889-1.578-0.9872.8942.6225.52944.116
1338488_A65A66:U93C94_88-0.446-0.6533.448-5.0583.52949.747-1.040.1423.4244.1775.98550.104
1338588_A66U67:A92U93_88-0.412-1.553.518-4.7153.18836.581-2.898-0.0253.4015.0417.45637.006
1225988_C45G46:C57G58_881.131-1.733.2682.7657.24227.72-5.025-1.6872.83114.754-5.63328.764
1226188_C47C62:U55G56_88-0.809-2.6033.38111.129.9543.832-4.1051.912.49912.892-14.40846.183
1339188_C115G116:C137G107_88-1.952-3.5623.37-1.985-1.2583.629-30.166-1.0164.761-17.63727.8144.323
1339388_C117C118:G135G136_88-0.961-2.4543.181-6.5378.37327.19-6.5170.6282.4917.0113.27929.154
1339488_C118C119:G134G135_88-0.627-1.9393.0622.7575.22629.902-4.6051.6752.62610.005-5.27830.467
1339588_C119C120:G133G134_88-0.01-2.0083.59-2.4649.24231.457-5.156-0.412.89216.5724.41932.844
1339688_C120U121:G132G133_880.114-1.8633.0551.9237.70537.538-3.6850.0382.63811.811-2.94838.339
1225388_G39A40:A105A104_88-2.276-3.3043.78210.161-12.0-131.7621.902-1.1633.716.5665.56-132.243
1225688_G42A43:U101U102_88-0.601-2.4382.977-14.2524.10230.562-4.782-1.0492.657.26925.25533.893
1226088_G46C47:G56C57_88-0.754-1.3583.189-3.1680.63138.502-2.130.7573.2170.9544.79438.632
1338188_G49C50:G75C76_880.011-1.1422.797-1.8543.20532.648-2.479-0.2862.6725.6793.28532.852
1338288_G63U64:A96A97_88-0.746-2.8033.68-11.416.79630.979-6.043-0.7233.0912.05320.23533.641
1338788_G68U69:A89C91_880.965-1.3083.0621.6262.39530.409-2.924-1.5332.9994.552-3.09130.543
1338988_G70A71:G87A88_880.238-1.8664.693.455-2.147-3.43438.0631.4782.11926.30842.332-5.323
1339088_G114C115:G107C108_88-0.25-2.2683.2170.9443.22336.365-4.0440.5243.0055.151-1.50936.514
1339288_G116C117:G136C137_880.671-1.4973.072-0.6874.55740.977-2.575-1.022.8866.4840.97741.224
1339988_G123G124:C129C130_880.12-2.0053.2123.1776.77831.59-4.6850.2992.73412.236-5.73532.443
1338388_U64A65:C94A96_883.938-2.0393.108-13.3613.16833.416-3.609-7.6911.2965.24222.11236.052
1338688_U67G68:C91A92_88-0.518-1.4573.3622.3716.08339.785-2.7971.0173.0818.866-3.45640.296
1338888_U69G70:A88A89_880.4611.1982.76-0.73511.80659.7660.685-0.4872.92311.7230.7360.819
1339788_U121U122:A131G132_88-0.023-2.353.123.58114.33825.556-6.990.6361.58629.48-7.36229.46
1339888_U122G123:C130A131_88-0.456-1.5473.269-1.493.22826.084-4.240.6123.0797.1093.28226.321

The above values were obtained using first alternate conformation only and calculated by 3DNA program.
Xiang-Jun Lu & Wilma K. Olson (2003). '3DNA: a software package for the analysis, rebuilding and visualization of three-dimensional nucleic acid structures', Nucleic Acids Res.31(17), 5108-21.
Xiang-Jun Lu & Wilma K. Olson (2008). '3DNA: a versatile, integrated software system for the analysis, rebuilding and visualization of three-dimensional nucleic-acid structures', Nat Protoc.3(7), 1213-27.