A Portal for Three-dimensional Structural Information about Nucleic Acids
As of 13-Oct-2021 number of released structures: 11721
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NDB ID: 4U4U  PDB ID: 4U4U 


Base Pair Morphology Step Parameters

CSV Format

Model NumberStep NumberStep NameShiftSlideRiseTiltRollTwistX-DisplacementY-DisplacementHelical RiseInclinationTipHelical Twist
14022_A40A41:U466G467_220.294-0.3643.76-2.49710.69760.123-0.941-0.4253.64110.5812.4761.026
14122_A41A438:G465U466_220.3580.2113.485-3.6565.48873.494-0.01-0.4223.4734.5773.0573.747
15822_A67G69:U82G83_220.825-0.3353.706-4.1043.85262.922-0.512-0.9923.6283.6823.92363.147
16022_A93A399:U395G396_22-2.014-0.0732.853-3.9974.293-1.535-11.593-30.477-0.553-55.143-51.346-6.063
17822_A103C309:G357U358_221.13-2.1833.171-6.0846.06192.974-1.607-0.8842.9974.1724.18893.279
17722_A105A103:U358C308_223.766-4.606-2.24542.53190.61916.904-0.471-3.616-2.9575.119-35.257101.142
18522_A108G109:C305U306_220.132-0.9922.849-6.5535.17232.89-2.404-1.1172.5948.95611.34733.904
18922_A112U113:A301U302_22-1.122-1.5092.965-1.78810.421.25-6.4892.2492.09126.2214.50823.699
19222_A119U120:U297C298_223.237-1.4993.15-8.8222.50152.41-1.816-4.1122.5442.8089.90353.15
19722_A124U125:G291U293_222.652-1.4214.203-8.73516.58743.807-3.299-4.1032.9621.1211.12247.467
19822_A138G142:A173A266_225.398-1.6924.283-9.2074.03977.243-1.486-4.6143.6443.2237.34677.793
110222_A145U146:A168A169_22-0.364-1.0773.221-13.485-4.06436.412-1.11-1.133.248-6.22420.65538.955
110422_A147A148:C166U167_221.157-2.043.95-0.98210.31647.212-3.413-1.5023.43512.7021.20948.272
110522_A148C149:G165C166_223.544-3.0183.581-5.86119.61544.283-4.959-4.6941.71824.5287.32948.569
111122_A182U183:A202U203_22-0.062-0.913.01-5.55714.00424.206-4.594-0.9612.13929.96811.89228.453
111722_A188C189:G196A197_220.565-0.8933.489-0.137-8.28958.966-0.446-0.5773.573-8.3760.13859.494
112222_A206U207:C258U259_220.442-1.5333.712-3.20719.90635.475-4.464-1.02.49629.8814.81440.643
112622_A210U211:A254U255_220.68-1.7883.4952.8045.37224.641-5.66-0.723.09812.351-6.44725.364
112922_A213G214:A251U252_22-0.722-0.8823.161-4.66917.17558.804-1.6370.52.87317.074.64161.208
113222_A236C237:G228U229_220.567-0.8573.397-2.95510.78549.98-1.746-0.8643.12612.5833.44851.137
114622_A265U267:A288U289_22-1.817-0.7732.769-3.8450.5743.592-1.0842.122.9010.7665.16543.757
18322_A312A315:A352A353_225.356-0.5792.8185.536-24.8325.0917.736-9.922.254-76.98-17.16325.938
115522_A316C317:G347U348_220.142-2.0843.1845.9345.7930.828-4.7620.7362.73710.652-10.91731.897
115822_A323U324:A344U345_22-0.961-1.1853.46-6.61821.39937.973-3.620.6582.59629.9259.25543.876
116322_A328G329:C339U340_220.232-1.8273.4112.2628.78226.71-5.7210.032.69518.351-4.72728.181
116622_A366A367:U374U375_22-0.074-1.5432.866-2.4143.32634.778-2.993-0.1882.715.5414.02135.013
116722_A367U368:G373U374_22-0.154-1.442.9212.5854.86739.366-2.6080.4862.7167.182-3.81439.735
16122_A399A401:C394U395_22-0.245-1.593.036-7.076-2.28338.372-2.117-0.4423.117-3.43410.64339.059
16222_A401G404:C393C394_223.52-1.4353.691.3841.48475.493-1.222-2.8273.721.212-1.1375.516
16622_A407C408:G389G390_22-1.3321.5332.7454.6468.293-14.611-9.039-2.1711.933-28.94516.217-17.417
16922_A410C411:G422G423_220.851-0.8692.929-10.1921.16326.793-1.977-3.6662.4112.421.04128.656
14222_A438U439:A464G465_22-3.41-3.682.718.5260.8018.336-10.47810.699-2.1372.852-65.93820.316
14422_A441C442:G462U463_22-0.843-2.6193.259-4.2756.41836.638-4.880.7842.85410.0736.70937.413
15622_A452G63:C87U88_22-2.813-1.7763.0243.6092.73855.106-2.0573.222.7652.954-3.89455.277
1122_A468G595:C31U32_221.705-2.793.742-8.118.97576.094-2.524-1.6253.287.2376.5476.905
117922_A485G486:U501U502_221.65-2.2494.57711.9857.78821.102-8.2851.1443.89418.745-28.84625.441
117422_A505U508:G480A481_220.037-4.3033.76526.218-4.45278.419-3.1420.763.829-3.432-20.20882.112
117822_A511A538:G540A477_22-2.734-4.536-3.589-30.03489.844168.177-2.1391.415-3.83945.06715.065172.0
118222_A515G516:C536G537_221.258-0.2573.825-3.5496.82857.924-0.677-1.5043.77.0163.64758.389
118422_A518A520:U532U533_22-2.133-1.3653.7514.08731.38945.36-3.1613.0281.83735.364-15.87156.378
118522_A520A521:C531U532_22-1.417-1.3733.8739.3239.91434.295-3.6233.6072.90916.051-15.09436.821
118622_A521U522:C530C531_22-0.231-1.633.773-0.58410.36449.484-2.7260.2253.39112.2260.68950.494
119222_A550G557:C587U588_22-1.46-0.7552.763-3.8733.70941.088-1.4071.7112.8085.2585.4941.422
1622_A599U600:A26U27_220.674-1.3552.973-0.290.26123.852-3.356-1.7162.950.6310.70223.855
1822_A601U602:G23U24_22-0.119-0.9473.2340.57611.61338.101-2.6720.2392.83517.304-0.85939.772
11122_A604C4:G20U21_22-3.164-0.7084.87728.383-3.32956.19-0.4844.7373.143-3.309-28.21262.508
120222_A615G616:C1105U1106_22-0.428-1.843.4712.74110.93531.919-4.8741.1642.66919.153-4.80133.802
120422_A622G1032:U1103U1104_220.591-3.2263.39811.5345.9886.53-17.316.4510.67129.229-56.29714.538
127222_A746C747:G802U805_224.024-1.4434.393-27.648-1.38661.618-1.246-4.8692.608-1.28625.65966.989
127922_A760G761:A789U790_22-0.573-0.7263.772.388-2.41763.906-0.5650.6593.772-2.281-2.25463.987
128122_A774G775:U785C786_22-0.123-3.925-3.255131.436-112.38-93.0381.018-1.169-4.05356.81966.454-175.136
128422_A814G816:A855U857_224.5670.8490.754-116.228111.932132.7120.398-2.316-0.37956.558.669172.553
114422_A844G845:C827U828_22-0.786-1.8223.6330.3824.12927.309-4.8911.7453.3158.683-0.80227.616
122222_A865G866:C962U965_223.103-1.5793.344.5362.68948.809-2.113-3.3773.5113.243-5.4749.076
122622_A869C870:G957U958_220.016-0.9163.224-1.5927.67946.453-1.755-0.1463.0429.6542.00247.074
123722_A881U882:U946U947_220.51-0.9252.807-1.775-3.00741.709-1.023-0.8752.841-4.2142.48841.848
124022_A884G885:C927U928_22-0.51-1.1273.5639.592-0.14434.787-1.7992.2833.316-0.236-15.67736.046
124322_A887U888:A924G925_22-0.129-1.0983.407-7.4084.8631.88-2.771-1.0563.168.64113.17333.057
124722_A891A892:U920U921_22-0.289-1.0893.234-2.2582.830.289-2.6150.113.1365.3354.30230.497
124822_A892U893:A919U920_220.647-1.3313.032-3.65515.77629.259-4.47-1.6261.98428.6346.63333.356
128822_A900G902:A907U909_22-1.13-0.9086.49410.121.34196.735-1.2761.0656.15614.121-6.68398.872
128922_A933U935:G876A944_222.6860.333-0.401-70.402151.31943.4410.575-1.161-0.92179.39336.939167.829
120922_A966A967:U632U633_22-1.585-2.2253.2390.0422.56135.455-4.0052.6023.0744.198-0.06935.544
121022_A967U968:G631U632_220.304-1.2563.2420.7774.86632.247-3.053-0.4113.0328.698-1.38832.612
121322_A970A971:G628U629_225.725-1.5513.103-1.024-0.83631.652-2.699-10.6632.96-1.5321.87631.679
121422_A971G972:C627G628_221.026-0.5793.750.66810.79263.933-1.057-0.9263.63210.116-0.62664.745
121622_A973A974:C625U626_22-0.04-1.5332.96-11.5094.56823.813-4.333-2.52.39510.2325.77926.799
121722_A974C975:G624C625_220.208-1.4283.2484.6472.77640.867-2.3240.1973.1523.954-6.6241.209
129322_A979G980:C1021C1022_220.061-1.1293.854-2.01310.2751.083-2.071-0.2243.57511.7682.30752.074
129722_A983G984:U1017U1018_220.17-2.1652.691-2.1818.43218.106-8.848-1.171.50925.0016.46720.076
130222_A992A993:G1011U1012_22-3.3030.5594.574-113.1-129.228-64.0280.871-2.6620.94266.074-57.828-172.99
130322_A993G994:C1010G1011_220.153-0.463.385-6.90813.42355.955-1.211-0.5373.17114.0247.21757.796
130522_A995U996:G1008U1009_220.621-1.5733.2820.142-1.08236.218-2.375-0.9793.328-1.739-0.22836.234
130722_A1036C1037:G1094U1095_220.843-1.9373.2076.0421.18832.357-3.62-0.4633.2362.108-10.72132.922
130922_A1061U1063:U1056U1058_226.144-3.261-0.63163.132135.062-58.0571.8493.078-0.009-74.6134.875-153.051
131122_A1065C1066:G1053U1054_221.163-2.0723.322-5.1694.79731.99-4.456-2.9042.7738.5749.23932.738
131522_A1069C1070:G1048U1049_220.519-1.5823.21-0.8084.99131.621-3.717-1.0792.9199.0861.47132.013
132222_A1076C1077:G1041G1042_221.155-1.2092.7351.2620.72925.089-2.958-2.3422.7531.676-2.90225.13
133222_A1113U1115:A1131A1132_22-3.2881.2930.223-125.229-126.06779.1140.8881.4041.221-63.3362.909178.222
133422_A1116U1117:U313G1130_22-3.622-1.9162.6390.3286.15135.772-3.7265.8522.2579.923-0.52936.281
11622_A1138U8:U15G16_22-0.6610.4132.3920.94-8.02413.6735.6132.8771.815-30.472-3.5715.871
12022_A1142A1143:A11U12_220.153-1.9583.5864.6297.44136.283-4.0890.43.13311.739-7.30337.292
12122_A1143U1144:G10A11_22-1.002-2.013.283-3.4413.76930.216-4.5351.2293.1087.1666.54330.634
12522_A1147C1148:G1629U1630_22-0.024-1.2723.1226.6849.78130.489-3.7851.0822.54317.779-12.14932.659
12822_A1151A1152:U1626U1627_22-1.546-1.4933.399-4.556-2.5735.333-2.0311.8063.66-4.2057.45635.706
12922_A1152G1153:C1625U1626_220.045-1.8773.2565.0668.43728.723-5.1480.852.58416.41-9.85330.329
13422_A1157A1160:U1617U1621_225.4910.0792.992-4.665-12.343114.3480.166-3.3042.835-7.3312.771114.837
13522_A1160C1161:G1616U1617_220.307-1.0533.8115.3824.10733.484-2.5440.4613.6647.04-9.22534.143
136122_A1203U1560:U1551G1553_221.4040.211-5.543-22.0210.871-53.677-0.1423.389-4.692-0.922-23.313-57.712
137522_A1211G1212:C1451U1452_221.101-1.73.6228.09113.76427.13-5.825-0.4562.68726.617-15.64631.404
138022_A1219C1220:G1263G1264_220.667-0.6433.515-0.741-5.03667.127-0.369-0.6333.543-4.5480.66967.297
138222_A1221C1222:G1261U1262_220.489-0.5753.141-4.8566.62234.643-1.844-1.4622.89310.9318.01535.574
138422_A1223A1224:U1259U1260_22-0.003-1.6923.2525.94119.41132.728-4.6740.6471.95631.014-9.49238.365
138822_A1234G1245:U1250U1251_222.25-2.368-0.453-100.746-140.182147.322-1.226-1.095-0.263-70.21150.46177.928
139022_A1236G1237:C1248U1249_221.014-1.4343.9223.9888.05930.412-4.357-1.0013.53714.956-7.40131.684
139222_A1238U1239:C1246U1247_222.121-1.812.88310.1967.44599.54-1.284-1.2412.934.862-6.658100.127
139922_A1275U1276:A1436C1431_220.619-3.7693.7132.9322.862-20.779-0.2252.0945.208-48.1166.167-30.938
141522_A1296G1297:A1300U1301_22-4.447-1.6783.1730.5059.86457.291-2.2134.6072.83810.204-0.52258.065
142322_A1336A1410:G1387U1415_225.0074.1023.77120.778-35.8283.5383.4631.1043.762-25.339-85.419126.008
142922_A1344A1345:U1380U1381_22-0.884-0.7053.0795.7333.67630.529-1.9392.6222.7696.875-10.72131.262
143022_A1345A1348:U1377U1380_221.9611.5734.60914.78913.404101.2080.718-0.9364.9178.614-9.504102.625
143122_A1348G1349:C1376U1377_22-0.056-1.2772.803-11.42110.77225.328-4.273-1.7831.96222.06923.39929.732
144022_A1357G1358:C1365U1366_22-0.144-1.4232.9673.73317.18428.576-4.7190.7391.81231.346-6.80933.457
144322_A1391U1392:A1406U1407_22-0.676-1.6453.4281.1239.03932.814-4.2241.332.86315.629-1.94234.021
142422_A1410C1338:G1386G1387_22-3.052-1.544-0.069124.924-123.736-143.4340.784-1.5140.27461.94262.536-178.692
135822_A1469C1470:G1574G1170_22-0.1060.3822.887-2.2732.21125.5320.306-0.332.9084.9795.11825.725
144822_A1471C1472:G1536U1538_221.229-2.7283.438-4.53111.227-80.0771.7810.8183.784-8.68-3.504-80.834
145122_A1475C1476:G1531U1532_220.677-1.3993.1616.0286.87741.463-2.597-0.3542.9699.574-8.39242.416
145522_A1479G1480:C1527U1528_22-1.042-2.2854.0149.94412.73327.709-6.5293.7522.27924.162-18.86831.995
145722_A1487G1488:C1518U1519_220.478-1.3592.909-11.852-5.74922.196-1.589-4.0792.594-13.51127.85425.769
137122_A1569A1570:G1541G1542_22-0.009-0.0252.632-7.8425.77230.123-0.906-1.1532.50810.75914.61631.623
137222_A1570C1571:G1540G1541_220.706-0.8173.572-5.589.43864.966-1.163-0.8933.3758.7225.15765.785
134022_A1576A1577:U1168G1169_221.365-1.5043.8580.9688.30455.859-2.129-1.383.648.81-1.02756.431
134122_A1577U1578:G1167U1168_220.505-1.3513.0365.497-1.34246.81-1.587-0.2083.109-1.682-6.88947.132
146722_A1586A1587:U1609G1610_22-0.237-0.5832.586-5.676-3.23760.392-0.4520.0152.622-3.2085.62460.711
146822_A1587G1588:U1608U1609_220.005-1.9043.8211.31917.73334.768-5.0950.162.58627.56-2.0538.926
147322_A1592A1593:U1603U1604_220.612-0.8953.504-2.5097.23346.58-1.738-0.9793.3039.0743.14847.17
147422_A1593G1594:C1602U1603_22-0.982-1.6533.1150.473.76928.928-4.0352.0432.8657.504-0.93529.171
147722_A1719G1720:A1681U1682_220.799-1.0212.8013.35218.599105.854-0.842-0.462.67311.596-2.09107.024
147922_A1721A1722:G1679G1680_22-0.65-0.3713.068-0.23814.18332.447-2.5381.0392.68424.0030.40335.335
148022_A1722U1723:A1678G1679_222.015-1.1833.267-1.1963.43353.408-1.526-2.3113.1493.8151.32953.522
148622_A1728C1729:G1672G1673_221.126-0.9733.204-6.636.6730.942-2.831-3.0992.6512.15412.08232.306
148822_A1730A1731:G1670A1671_220.74-2.1313.1187.8951.775-23.3814.473.9912.869-4.22118.775-24.723
148922_A1731A1732:U1669G1670_220.527-0.7493.481-5.8134.32157.986-1.006-0.8593.364.4385.97158.398
149022_A1732C1733:G1668U1669_22-0.592-1.1713.2942.7685.56738.846-2.3861.23.0568.305-4.1339.321
149822_A1740U1741:A1660U1661_220.513-1.5223.0352.3430.47630.893-2.934-0.543.0410.892-4.3930.983
150222_A1744G1745:A1655U1656_22-1.975-0.4233.6284.35918.0757.426-1.3832.2033.23218.287-4.41160.115
150422_A1746G1747:C1653G1654_221.553-0.1954.165-5.89215.15663.528-0.965-1.7433.89514.1525.50265.364
150722_A1749A1750:U1650A1651_22-0.773-1.9213.3581.3244.95618.695-8.0292.9152.70114.896-3.97819.38
150822_A1750C1751:G1649U1650_220.812-1.3462.9280.8552.58330.737-2.972-1.382.8294.862-1.60830.855
151122_A1753A1754:C1646U1647_22-0.761-3.341-0.775-150.30890.04695.653-1.4670.722-1.28945.34175.685176.789
151222_A1754A1756:G1645C1646_22-2.979-2.51.80286.709-103.926162.826-1.1561.5721.786-52.137-43.499173.497
151322_A1756G1757:C1644G1645_22-0.114-2.2193.694-8.95821.02839.568-4.698-0.632.27228.37912.08945.465
151822_A1762A1763:C1639C1640_22-1.292-1.0283.327-7.741-5.05641.095-0.8710.9413.598-7.09510.86242.078
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1337288_G68U69:A89C91_880.9-1.3182.928-2.7115.52230.198-3.394-2.1422.56410.4635.13630.804
1337488_G70A71:G87A88_880.249-1.7634.635-1.482-7.485-2.6535.142-2.711-0.06669.524-13.766-8.077
1337588_G114C115:G107C108_88-0.12-2.433.229-0.9292.50236.245-4.2330.0683.0624.0161.49136.34
1337788_G116C117:G136C137_880.259-1.5143.034-6.4224.25439.949-2.604-1.0132.7926.1579.29540.656
1338488_G123G124:C129C130_88-0.361-1.763.66711.22414.79336.705-4.1521.7592.58921.859-16.58640.99
1226685_U26C339:G25G337_850.2443.2510.58-152.0378.343-60.833-1.60.179-0.802-40.184-77.981-172.265
1226588_U27U26:G337A336_55-0.5671.458-2.417-7.047-1.309-30.379-2.897-0.049-2.422.456-13.218-31.193
1226388_U29C28:G335A334_55-0.7731.527-3.01-3.442-2.333-35.932-2.762-0.801-2.9673.766-5.557-36.164
1336888_U64A65:C94A96_884.235-2.0192.854-16.2260.88635.097-3.122-7.6010.8351.38125.338.569
1337188_U67G68:C91A92_88-0.642-1.413.3274.1185.70838.133-2.8081.463.018.646-6.23838.754
1337388_U69G70:A88A89_88-0.0821.2012.903-0.36113.35156.2610.5770.0673.09213.9410.37757.698
1338288_U121U122:A131G132_880.119-2.2812.9593.1299.95721.372-8.0270.4911.73625.016-7.86223.758
1338388_U122G123:C130A131_88-0.218-1.6213.383-2.9332.12627.854-3.849-0.2533.2584.3926.0628.084

The above values were obtained using first alternate conformation only and calculated by 3DNA program.
Xiang-Jun Lu & Wilma K. Olson (2003). '3DNA: a software package for the analysis, rebuilding and visualization of three-dimensional nucleic acid structures', Nucleic Acids Res.31(17), 5108-21.
Xiang-Jun Lu & Wilma K. Olson (2008). '3DNA: a versatile, integrated software system for the analysis, rebuilding and visualization of three-dimensional nucleic-acid structures', Nat Protoc.3(7), 1213-27.